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Richard Nash

Telefon: 480 36 416
Besøksadresse: Institute of Marine Research Nordnesgaten 33/PB 1870 Nordnes 5817 Bergen Norway Office phone: +47 48036416

Arbeider med

Marine Ecology (fish and fisheries) – have worked in many different areas of marine biology (including a short stop over in Limnology – Lake Michigan). Started with the behavioural ecology of small demersal burrowing fish in British waters. Have worked with fish community ecology in the Oslofjord and the Azores, early life history dynamics in the Great Lakes, distribution of organisms using traditional capture techniques and acoustics on the north wall of the Gulf Stream, acoustic assessments of herring in the Gulf of Maine and UK waters, flatfish nursery ground dynamics in UK waters and stock discrimination techniques again in UK waters.

Currently my research covers the following areas:
• The dynamics of flatfish nursery grounds
• Stock discrimination for herring
• Early life history dynamics of commercially exploited fish stocks
• Stock Reproductive Potential of cod and herring.


Fish life-history dynamics; population ecology; stock reproductive potential; recruitment processes; Irish sea; North Sea; Barents/Norwegian Seas; plaice; herring; cod; non-commercial species.

Curriculum Vitae


BSc (Marine Biology) University College Swansea, UK, 1977
PhD (Fish ecology) University of Glasgow, UK. University Marine Biological Station and Dunstaffnage Marine Laboratory (CASE institution), 1980


1981 82 Post-doctoral researcher. Royal Society of London, European Science Exchange Programme, to University of Oslo, Norway. Research: The effects of eutrophication on demersal fish communities in the Oslofjord.
1982 84 Post-doctoral researcher. Wisconsin Sea Grant, University of Wisconsin-Madison, USA. Research: Species interactions and early life history dynamics of forage fish in Lake Michigan.
1983 86 Post-doctoral researcher. Office of Naval Research, University of Wisconsin-Madison, USA. Research: Acoustical techniques for the study of nekton and zooplankton at water type boundaries and ocean fronts.
1986 87 Faculty Associate - University of Maine, Orono, USA.
1986-90 Partner - Mainely Fish, Research, Searsport, Maine, USA.
1987 Adjunct Faculty - Unity College, Unity, Maine, USA.
1988-00 Lecturer - Port Erin Marine Laboratory, University of Liverpool, UK.
2000-04 Senior Lecturer - Port Erin Marine Laboratory, University of Liverpool, UK.
2003-04 Sabbatical year at the Institute for Fisheries and Marine Biology (Department of Biology), University of Bergen, Norway.
2005- Institute of Marine Research, Bergen

NMA Advanced Course:

Course leader. Nursery ground ecology: Methods of study and analysis. Bergen, Norway. 1-10th August 2009.

Recent publications (2005 to present):

Nash, R.D.M. & Geffen, A.J. 2005. Chapter 7. Age and growth. Pp. 138-163. In R.N. Gibson (ed) Flatfishes: Biology and Exploitation. Blackwell Science Ltd, Oxford.

Nash, R.D.M. & Geffen, A.J. 2005. Aspects of the general biology of Sars eelpout (Lycenchelys sarsi) in the Oslofjord, Norway. Marine Biology Research 1: 33-38.

Nash, R.D.M. & Dickey-Collas, M. 2005. The influence of life history dynamics and environment on the determination of year class strength in North Sea herring (Clupea harengus L.). Fisheries Oceanography 14: 279-291.

Lee, O., Nash, R.D.M. & Danilowicz, B. 2005. Small-scale spatiotemporal variability in ichthyoplankton and zooplankton distribution in relation to a tidal-mixing front in the northwestern Irish Sea. ICES Journal of Marine Science 62: 1021-1036.

Figueiredo, G.M. de, Nash, R.D.M. & Montagnes, D.J.S. 2005. The role of the generally unrecognised microprey source as food for larval fish in the Irish Sea. Marine Biology 148: 395-404.

Nash, R.D.M., Valencia, A.H. & Geffen, A.J. 2006. The origin of Fulton’s Condition Factor – setting the record straight. Fisheries 31: 236-238.

Fox, C.J., McCloghrie, P., Young, E.F. & Nash, R.D.M. 2006. The importance of individual behaviour for successful settlement in juvenile plaice - a modelling and field study in the eastern Irish Sea. Fisheries Oceanography 15: 301-313.

Geffen, A.J., Fox, C.J. & Nash, R.D.M. 2006. Temperature dependent development rates of cod (Gadus morhua L.) eggs. Journal of Fish Biology 69: 1060-1080.

Rowlands, W. Ll., Dickey-Collas, M., Geffen, A.J. & Nash, R.D.M. 2006. Gape morphology of western Irish Sea cod (Gadus morhua L.), haddock (Melanogrammus aeglefinus L.) and whiting (Merlangius merlangus L.) through larval development to the juvenile stage. Journal of Fish Biology 69: 1379-1395.

Kennedy, J., Witthames, P.R. & Nash, R.D.M. 2007. The concept of fecundity regulation in plaice (Pleuronectes platessa) tested on three Irish Sea spawning populations. Canadian Journal of Fisheries and Aquatic Sciences 64: 587-601.

Geffen, A.J., van der Veer, H.W. & Nash, R.D.M. 2007. The cost of metamorphosis in flatfishes. Journal of Sea Research 58: 35-45. 

Kennedy, J., Nash, R.D.M. & Geffen, A.J. 2007. Maternal effects on egg and larval characteristics of the European plaice (Pleuronectes platessa). Journal of Sea Research 58: 65-77.

Beggs, S.E. & Nash, R.D.M. 2007. Variability in settlement and recruitment of O-group dab Limanda limanda L. in Port Erin Bay, Irish Sea. Journal of Sea Research 58: 90-99

Figueiredo, G.M. de, Nash, R.D.M. & Montagnes, D.J.S. 2007. Do protozoa contribute significantly to the diet of larval fish in the Irish Sea? Journal of the Marine Biological Association, UK. 87: 843-850.

Nash, R.D.M., Geffen, A.J., Burrows, M.T. & Gibson, R.N. 2007. The dynamics of shallow-water juvenile flatfish nursery grounds: application of the ‘self-thinning rule’. Marine Ecology Progress Series 344: 231-244.

Fox, C.J., Geffen, A.J., Taylor, N., Davison, P., Rossetti, H. & Nash, R.D.M. 2007. Birth-date selection in early life stages of plaice Pleuronectes platessa in the eastern Irish Sea (British Isles). Marine Ecology Progress Series 345: 255-269.

Lee, O., Nash, R.D.M. & Danilowicz, B. 2007. Small-scale variability in growth and condition of dab (Limanda limanda) larvae in relation to a tidal-mixing front off the west coast of the Isle of Man. Journal of Fish Biology 71: 1056-1068.

Morato, T., Afonso, P., Carvalho, N., Lourinho, P., Santos, R.S., Krug, H.M. & Nash, R.D.M. 2007. Growth, reproduction and recruitment patterns of the wide-eyed flounder, Bothus podas Delaroche (Pisces: Bothidae), from the Azores. Marine Biology Research 3: 403-411.

Kennedy, J., Witthames, P.R., Nash, R.D.M. & Fox. C.J. 2008. Is fecundity in plaice (Pleuronectes platessa L.) down regulated in response to reduced food intake during autumn? Journal of Fish Biology 72: 78-92.

Watts, P.C., O’Leary, D., Cross, M.C., Coughlan, J., Dillane, E., Kay, S.M., Wylde, S., Stet, R., Nash, R.D.M., Hatfield, E.M.C. & Cross, T.F. 2008. Contrasting levels of genetic differentiation among putative neutral microsatellite loci in Atlantic herring Clupea harengus populations and the implications for assessing stock structure. Hydrobiologia 606: 27-33.

Rowlands, W. Ll., Dickey-Collas, M., Geffen, A.J. & Nash, R.D.M. 2008. Diet overlap and prey selection through metamorphosis in Irish Sea cod (Gadus morhua), haddock (Melanogrammus aeglefinus) and whiting (Merlangius merlangus). Canadian Journal of Fisheries and Aquatic Sciences 65: 1297-1306. 

Nash, R.D.M., Kjesbu, O.S., Trippel, E.A., Finden, H. & Geffen, A.J. 2008. The potential importance of the paternal contribution to the ‘Stock Reproductive Potential’ of North-East Arctic cod (Gadus morhua). Journal of Northwest Atlantic Fishery Science 41: 71-83. 

Nash, R.D.M., Dickey-Collas, M. & Kell, L.T. 2009. Stock and recruitment in North Sea herring (Clupea harengus); compensation and depensation in the population dynamics. Fisheries Research 95: 88-97.

Fox, C.J., McCloghrie, P. & Nash, R.D.M. 2009. Potential transport of plaice eggs and larvae between two apparently self-contained sub-populations in the Irish Sea. Estuarine, Coastal and Shelf Science 81: 381-389.

Payne, M.R., Hatfield, E.M.C., Dickey-Collas, M., Falkenhaug, T., Gallego, A., Gröger, J., Licandro, P., Llope, M., Munk, P., Röckmann, C., Schmidt, J.O. & Nash, R.D.M. 2009. Recruitment in a changing environment: the 2000s North Sea herring recruitment failure. ICES Journal of Marine Science 66: 272-277. 

Payne, M.R., Dickey-Collas, M. & Nash, R.D.M. 2009. Comment on “Does the fall phytoplankton bloom control recruitment of Georges Bank haddock, Melanogrammus aeglefinus, through parental condition?”. Canadian Journal of Fisheries and Aquatic Sciences 66: 869–872. 

Figueiredo, G.M. de, Montagnes, D.J.S. & Nash, R.D.M. 2009. The importance of protozooplankton as prey for copepods in the coastal areas of the central Irish Sea. Hydrobiologia 628: 227-239.

Oeberst, R., Klenz, B., Gröhsler, T., Dickey-Collas, M., Nash, R.D.M. & Zimmermann, C. 2009. When is year-class strength determined in western Baltic herring?. ICES Journal of Marine Science 66: 1667-1672.

Oeberst, R., Dickey-Collas, M. & Nash, R.D.M. 2009. Mean daily growth of herring larvae in relation to temperature over a range of 5–20oC, based on weekly repeated cruises in the Greifswalder Bodden. ICES Journal of Marine Science 66: 1696–1701. 

Kell, L.T., Dickey-Collas, M., Hintzen, N.T., Nash, R.D.M., Pilling, G.M. & Roel, B.A. 2009. Lumpers or splitters? evaluating recovery and management plans for metapopulations of herring. ICES Journal of Marine Science 66: 1776–1783.

Kennedy, J., Skjæraasen, J.E., Nash, R.D.M., Thorsen, A., Slotte, A., Hansen, T. & Kjesbu, O.S. 2010. Do capital breeders like Atlantic herring (Clupea harengus L.) exhibit sensitive periods of nutritional control on ovary development and fecundity regulation? Canadian Journal of Fisheries and Aquatic Sciences 67: 16-27.

Skjæraasen, J.E., Nash, R.D.M., Kennedy, J., Thorsen, A., Nilsen, T. & Kjesbu, O.S. 2010. Liver energy, atresia and oocyte stage influence fecundity regulation in Northeast Arctic cod. Marine Ecology Progress Series 404: 173-183.

Watts, P.C., Kay, S.M., Wolfenden, D., Fox, C.J., Geffen, A.J., Kemp, S.J. & Nash, R.D.M. 2010. Temporal patterns of fine-scale spatial genetic structure and effective population size in European plaice (Pleuronectes platessa L.) along the west coast of Scotland and the Irish Sea. ICES Journal of Marine Science 67: 607-616. 

Ndjaula, H.O.N., Nash, R.D.M., Slotte, A., Johannessen, A. & Kjesbu, O.S. 2010. Long-term changes in the total egg production of Norwegian spring-spawning herring Clupea harengus (L.) – implications of variations in population structure and condition factor. Fisheries Research 104: 19-26.

Thorsen, A., Witthames, P.R., Marteinsdottir, G., Nash R.D.M. & Kjesbu, O.S. 2010. Comparing fecundity of various cod (Gadus morhua) stocks in the northern Atlantic in a standardized way. Fisheries Research 104: 45-55.

 Nash, R.D.M., Pilling, G.M., Kell, L.T., Schön, P-J. & Kjesbu, O.S. 2010. Investment in maturity at age and length in northeast Atlantic cod stocks. Fisheries Research 104: 89-99.

Dickey-Collas, M., Nash, R.D.M., Brunel, T., van Damme, C.J.G., Marshall, C.T., Payne, M.R., Corten, A., Geffen, A.J., Peck, M.A., Hatfield, E.M.C., Hintzen, N.T., Enberg, K., Kell, L.T. & Simmonds, E.J. 2010. Lessons learned from stock collapse and recovery of North Sea herring: a review. ICES Journal of Marine Science 67: 1875–1886.

Ciotti, B., Targett, T., Nash, R.D.M., Batty, R.S., Burrows, M.T. & Geffen, A.J. 2010. Development, validation and field application of an RNA-based growth index in juvenile plaice Pleuronectes platessa. Journal of Fish Biology 77: 2181–2209.

Geffen, A.J., Nash, R.D.M., Dau, K. & Harwood, A.J.P. 2011. Sub-cohort dynamics of O-group plaice, Pleuronectes platessa L., in the Northern Irish Sea: settlement, growth and mortality. Journal of Experimental Marine Biology and Ecology.

Kennedy, J., Skjæraasen, J.E., Nash, R.D.M., Slotte, A., Geffen, A.J. & Kjesbu, O.S. 2011. Evaluation of the frequency of skipped spawning in Norwegian spring-spawning herring. Journal of Sea Research.

Geffen, A.J. & Nash, R.D.M. 2011. Egg development rates for use in egg production methods (EPMs) and beyond. Fisheries Research.

Kennedy, J., Nash, R.D.M., Slotte, A. & Kjesbu, O.S. 2011. The reproductive strategy in the determination of fecundity and abortive maturation in Norwegian Spring-spawning herring (Clupea harengus). Marine Biology.