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Halvor Knutsen

Phone: 924 09 667
Visiting address: Havforskningsinstituttet Forskningsstasjonen Flødevigen Nye Flødevigveien 4817 HIS

Works with

My primary research interest is on connectivity and microevolutionary patterns and processes. I started working on such issues in 2000. Since then, I have worked on linking genetic data with physical and biological data in different modeling approaches in studies of several high gene flow species.The novel outcome of these papers was a better understanding of underlying mechanisms behind the observed genetic patterns; for instance, determining during which lifestage gene flow more likely occurs. Another outcome was knowledge on the extent these spatial differences are caused by ecological responses (i.e. plasticity) or local adaptations. I  have also investigated mechanisms behind genetic patterns by ecological studies of egg density and current measurements in fjords, or behaviour data, all combined with advanced statistics.

I have been involved in and also lead (as PI) several research projects that have received funding from several sources in EU, ESF (European Science Foundation), the Norwegian Research Council (NRC), and Nordic funding sources (Nordic Council). These projects have allowed me to stay active as a researcher and to publish within my preferred fields of research

Curriculum Vitae

Barceló Caren, Ciannelli Lorenzo, Olsen Esben Moland, Johannessen Tore, Knutsen Halvor, Eight decades of sampling reveal a contemporary novel fish assemblage in coastal nursery habitat (2015). Global Change Biology . ISSN 1354-1013. doi: 10.1111/gcb.13047

Quintela María, Danielsen Elin A., Lòpez Lùa, Barreiro Rodolfo, Svåsand Terje, Knutsen Halvor, Skiftesvik Anne Berit, Glover Kevin, Genetic analysis of ballan wrasse from Galicia (NW Spain) demonstrates within-species genetic divergence associated with plain and spotted morphotype frequencies (2015). Integrative Zoology . ISSN 1749-4877. doi: in press

Roney Nancy E., Hutchings Jeffrey, Olsen Esben Moland, Knutsen Halvor, Albretsen Jon, Kuparinen Anna, Fine-scale life-history structure in a highly mobile marine fish. (2015). Evolutionary Ecology Research . ISSN 1522-0613.

Ellis Charlie, Hodgson DJ, Andre Carl, Knutsen Tonje, Knutsen Halvor, Griffiths Amber, Genotype Reconstruction of Paternity in European Lobsters (Homarus gammarus) (2015). PLoS ONE . ISSN 1932-6203.

Freitas Carla, Olsen Esben Moland, Knutsen Halvor, Albretsen Jon, Moland Even, Temperature-associated habitat selection in a cold-water marine fish. (2015). Journal of Animal Ecology . ISSN 0021-8790. s - doi: 10.1111/1365-2656.12458

Catarino Diana, Knutsen Halvor, Verissimo Ana, Olsen Esben Moland, Jorde Per Erik, Menezes Gui, Sannæs Hanne, Stankovi? David, company Joan, Neat Francis, Danovaro Roberto, Dell'Anno Antonio, Rochowski Bastien, Stefanni Sergio, The Pillars of Hercules as a bathymetric barrier to gene-flow promoting isolation in a global deep-sea shark (Centroscymnus coelolepis) (2015). Molecular Ecology . ISSN 0962-1083. doi: DOI: 10.1111/mec.13453

Freitas Carla, Olsen Esben Moland, Moland Even, Ciannelli Lorenzo, Knutsen Halvor, Behavioral responses of Atlantic cod to sea temperature changes (2015). Ecology and Evolution . ISSN 2045-7758. 5 (10), s 2070 - 2083 doi: 10.1002/ece3.1496

Berg Paul Ragnar, Jentoft Sissel, Star Bastiaan, Ring Kristoffer Hofaker, Knutsen Halvor, Lien Sigbjørn, Jakobsen Kjetill Sigurd, Andre Carl, Adaptation to Low Salinity Promotes Genomic Divergence in Atlantic Cod (Gadus morhua L.) (2015). Genome Biology and Evolution . ISSN 1759-6653. 7 (6), s 1644 - 1663 doi: 10.1093/gbe/evv093

Blanco Gonzalez Enrique, Aritaki Masato, Knutsen Halvor, Taniguchi Nobuhiko, Effects of large-scale releases on the genetic structure of red sea bream (Pagrus major, Temminck et Schlegel) populations in Japan (2015). PLoS ONE . ISSN 1932-6203. 10:e0125743 (5), s - doi: 10.1371/journal.pone.0125743

Rogers Lauren, Olsen Esben Moland, Knutsen Halvor, Stenseth Nils Christian, Habitat effects on population connectivity in a coastal seascape (2014). Marine Ecology Progress Series . ISSN 0171-8630. 511 s 153 - 163 doi: 10.3354/meps10944

Knutsen Halvor, Jorde Per Erik, Blanco Enrique, Ole Eigaard, Pereyra Ricardo T., Sannæs Hanne, Dahl Mikael, Andre Carl, Søvik Guldborg, Does population genetic structure support present management regulations of the northern shrimp (Pandalus borealis) in Skagerrak and the North Sea? (2015). ICES Journal of Marine Science . ISSN 1054-3139. s - doi: 10.1093/icesjms/fsu204

Longmore Craig, Trueman Clive N., Neat Francis, Jorde Per Erik, Knutsen Halvor, Stefanni Sergio, Catarino Diana, Milton James A., Mariani Stefano, Ocean-scale connectivity and life cycle reconstruction in a deep-sea fish (2014). Canadian Journal of Fisheries and Aquatic Sciences . ISSN 0706-652X. 71 (9), s 1312 - 1323 doi: 10.1139/cjfas-2013-0343

Blanco Gonzalez Enrique, Knutsen Halvor, Jorde Per Erik, Glover Kevin, Bergstad Odd Aksel, Genetic analyses of ling (Molva molva) in the Northeast Atlantic reveal patterns relevant to stock assessments and management advice (2015). ICES Journal of Marine Science . ISSN 1054-3139. 72 (2), s 635 - 641 doi: 10.1093/icesjms/fsu135

Quintela, María; Elin, Danielsen; Svåsand, Terje; Knutsen, Halvor; Skiftesvik, Anne Berit; Glover, Kevin (2014). Isolation and characterization of twenty microsatellite loci for the ballan wrasse, Labrus bergylta. Conservation Genetics Resources. In press

Huserbråten BO, Moland E, Knutsen H, Olsen EM, André C, Stenseth NC (2013) Conservation, spillover and gene flow within a network of northern European marine protected areas. PLOS ONE, doi: 10.1371/journal.pone.0073388

Knutsen, Halvor; Jorde, Per E., Blanco Enrique G., Robalo, Joana I. Almada, V (2013). Climate change and genetic structure of leading edge and rear end populations in a northwards shifting marine fish species, the corkwing wrasse (Symphodus melops). PLOS ONE, doi: 10.1371/journal.pone.0067492

Moland, Even; Olsen, Esben Moland; Knutsen, Halvor; Garrigou, Pauline; Espeland, Sigurd Heiberg; Kleiven, Alf Ring; Andre, Carl & Knutsen, Jan Atle (2013). Lobster and cod benefit from small-scale northern marine protected areas: inference from an empirical before - after control-impact study. Proceedings of the Royal Society of London. Biological Sciences.  ISSN 0962-8452.  280(1754) . doi: 10.1098/rspb.2012.2679

Knutsen, Halvor; Jorde, Per Erik; Bergstad, Odd Aksel & Skogen, Morten D. (2012). Population genetic structure in a deepwater fish Coryphaenoides rupestris: patterns and processes . Marine Ecology Progress Series.  ISSN 0171-8630.  460, s 233- 246 . doi: 10.3354/meps09728

Pereyra, Ricardo T.; Westgaard, Jon-Ivar; Dahl, Mikael; Johansen, Torild; Knutsen, Halvor; Ring, Anna-Karin; Søvik, Guldborg & André, Carl (2012). Isolation and characterization of nuclear microsatellite loci in the northern shrimp, Pandalus borealis.. Conservation Genetics Resources.  ISSN 1877-7252.  4(1), s 109- 112 . doi: 10.1007/s12686-011-9486-4

Robalo, Joana I.; Castilho, Rita; Francisco, Sara M.; Almada, Frederico; Knutsen, Halvor; Jorde, Per Erik; Pereira, Ana M. & Almada, Vitor C. (2012). Northern refugia and recent expansion in the North Sea: the case of the wrasse Symphodus melops (Linnaeus, 1758). Ecology and Evolution.  ISSN 2045-7758.  2(1), s 153- 164 . doi: 10.1002/ece3.77

Knutsen H, Olsen EM, Jorde PE, Espeland SH, André C, Stenseth NC (2011). 'Are Low but Statistically Significant Levels of Genetic Differentiation in Marine Fishes “Biologically Meaningful"? Molecular Ecology. 20, 768-783

Rogers, L, Stige LC, Olsen EM, Knutsen H, Sik-Chan K, Stenseth N.Chr. (2011). Cod phenotypes respond to changing climate and density throughout a century on the Norwegian coast. PNAS, 108, 1961-1966.

Ciannelli L., Knutsen H., Olsen E. M., Espeland S., Asplin L., Jelmert A., Knutsen J. A., Stenseth N Chr. 2010. Maintainance of small-scale genetic structure in a marine population in relation to water circulation and egg characteristics. Ecology, doi:10.1890/09-1548

Espeland SH, Thoresen AG, Olsen EM, Stige LC, Knutsen H, Gjøsæter J, Stenseth NC 2010. Diel vertical migration patterns in young Skagerrak coastal cod. MEPS, In Press

Knutsen J. A., Knutsen H., Rinde E., Christie H., Bodvin T., Dahl E. 2010. Mapping Biological Resources in the Coastal Zone – an Evaluation of Methods in a Pioneering Study from Norway. Ambio, DOI: 10.1007/s13280-010-0023-6

André C, Knutsen H., 2010. Development of twelve novel microsatellite loci in the European lobster (Homarus gammarus) .Conservation Genetic Resources, DOI 10.1007/s12686-009-9151-3

Glover K, Dahle G, Westgaard JI, Johansen T, Knutsen H and K. E. Jørstad 2010.Genetic diversity within and among Atlantic cod (Gadus morhua) farmed in marine cages: a proof-of-concept study for the identification of escapees. Animal genetics, doi:10.1111/j.1365-2052.2010.02025.x.

Knutsen H, Sannæs H. 2009, Development of twelve microsatellite loci in the corkwing wrasse (Symphodus melops). Conservation Genetic Resources, DOI 10.1007/s12686-009-9100-1

Knutsen H., Catarino D., Sannæs H., Stefanni S., 2009, Development of eleven microsatellite loci in the deep sea black scabbardfish (Aphanopus carbo).
Conservation Genetic Resources, Conservation Genetic Resources, 1, 89-92.

Espeland S.H., Olsen M. E., Knutsen H., Gjøsæter J., Danielssen D., Stenseth N. C., 2009, New perspectives on fish movement: kernel and GAM smoothers applied to a century of tagging data on coastal cod. MEPS, 372, 231-241

Knutsen H, Jorde PE, Sannæs H, Hoelzel AR, Bergstad OA, Stefanni S, Johansen T and Stenseth NC, 2009, Bathymetric barriers promoting genetic structure in the deepwater demersal fish tusk Brosme brosme. Molecular Ecology, in press

Ring AK, Knutsen H and André C, 2009, 'Development of ten microsatellite loci in the ling (Molva molva)
Molecular Ecology Resources, in press

Schneider M, Sannæs H, Jorde, PEJ and Knutsen, H, 2009, Isolation and characterisation of 11 microsatellite loci in the abbysal carapine grenadier Coryphanoides carapipnus (Actinoperygii, Macrouridae) and cross amplification in two other deep-sea macrourid species
Conservation genetics, DOI 10.1007/s10592-009-9838-8

Stefanni S, Bettencourt R, Knutsen H and Menezes G, 2009, Rapid PCR-RFLP method for discrimination of the two Atlantic cryptic deep-sea species of scabbardfish
Molecular Ecology Resources, doi: 10.1111/j.1755-0998.2008.02470.

Knutsen, H., Marie Le Goff-Vitry, Fiani, D., Hoelzel, A. R, 2008, Isolation and characterization of microsatellite loci in a marine fish species, the roundnose grenadier (Coryphanoides rupestris)
Molecular Ecology Notes, 8, 993-995

Olsen EM, Knutsen H, Gjøsæter J, Jorde PE, Knutsen JA and Stenseth NC, 2008, Small-scale biocomplexity in coastal Atlantic cod supports a Darwinian perspective on fisheries management
Evolutionary Applications, doi:10.1111/j.1752-4571.2008.00024.x

Knutsen H., Olsen, E.M., Ciannelli, L., Espeland, S., Knutsen, J.A., Simonsen J.H., Skreslet, S., Stenseth, N.Chr, 2007, Egg distribution, bottom topography and small-scale population structure in a coastal marine system
MEPS, 333, 249-255

Knutsen, H., Jorde, P.E., Albert, O.T.,Hoelzel, A.R. & Stenseth, N.Chr., 2007, Population genetic structure influenced by oceanic current systems in the North Atlantic Greenland halibut
CJFAS, 64, 857-866

Stefanni, S.& Knutsen H., 2007, Phylogeography and demographic history of the deep-sea fish, Aphanopus carbo, in the NE Atlantic: vicariance followed by secondary contact or speciation?
Molecular Phylogony and Evolution, 42, 38–46

Knutsen, H., Fiani, D., Sannæs, H., Hoelzel, A. R., 2007, Isolation and characterization of microsatellite loci in a marine fish species, the tusk (Brosme brosme)
Mol. Ecol. notes,7, 851-853

Sigurd Heiberg Espeland, Ailin Fernløf Gundersen, Esben Moland Olsen, Halvor Knutsen, Jakob Gjøsæter and Nils Chr. Stenseth, 2007, The Maintenance of small-scale population structure in coastal cod (Gadus morhua)
ICES J. M. Sci, 64, 920-928

Jorde, P. E., Knutsen, H. and Stenseth, N. C. , 2007, Population structuring of coastal cod (Gadus morhua L.) and the geographic extent of local populations
MEPS, 343,229-237.

Knutsen, J. A., Knutsen, H., Olsen, E. M., Jonsson, B. , 2004, Marine feeding of anadromous brown trout (Salmo trutta L.) during winter.
Journal of Fish Biology, 64, 1-11.

Stenseth, N.C., Jorde, P. E., Chan, K.-S., Hansen, Knutsen H., Andre, C., Skogen, M.D. & Lekve, K., 2006, Ecological and genetic impact of larval drift: the Atlantic cod
Proceedings of Royal Society Series B, B 273 (1590): 1085-1092)

Knutsen, H., Jorde, P. E., André, C. and Stenseth, N. C. , 2003, Fine-scaled geographic population structuring in a highly mobile marine species: the Atlantic cod.
Molecular Ecology 12, 385-394.
Knutsen et al. 2003.pdf

Knutsen, J.A.K, Knutsen H., Paulsen, Ø., Kristensen, Ø., Vike, S. , 2003, Biologiske verdier i sjø i Tvedestrand kommune.
Fisken og Havet, 7, 66s.

Esben Moland Olsen, Halvor Knutsen, Jan Henrik Simonsen; Bror Jonsson; Jan Atle Knutsen, 2006, Seasonal progression of marine growth
Ecology of Freshwater Fish, 15, 446-452

Knutsen, H., André, C., Jorde, P.E., , Skogen, M.D., Thuróczy, E. & Stenseth N.C.S, 2004, Transport of North Sea cod larva into the Skagerrak coastal populations
Proc. R. Soc. Lond., BB 271, 1337-1344

Knutsen, J.A., Knutsen, H., Gjøsæter, J & Jonsson, B., 2001, Feeding of anadromous brown trout at sea
Journal of Fish Biology, 59, 533-543

Heuch, P. A., Knutsen, J. A., Knutsen, H. and Schram, T. A., 2002, Sea lice (Lepeophtheirus salmonis and Caligus elongatus) over-wintering on sea trout (Salmo trutta) in coastal areas of the Skagerrak.
J.Mar.Biol.Ass., 82, 1-6.

Knutsen, H, Knutsen, J. A., Jorde, P. E. , 2001, Genetic evidence for mixed origin of recolonized sea-trout populations.
Heredity, 87, 1-8.

Knutsen, H., Rukke, B. A., Jorde, P. E. and Ims, R. A. , 2000, Genetic differentiation among populations of the beetle Bolitophagus reticulatus (Coleoptera: Tenebrionidae) in a fragmented and a continuous landscape.
Heredity, 6, 767-776

Olsen,E. M., Knutsen, H., Stenseth, N. Chr., Gjøsæter, J. and Jorde,P. E. , 2004, Life-history variation among genetically different neighbouring populations of coastal Atlantic cod
Journal of Fish Biology, 64, 1-6.

Knutsen H., Jorde, P. E. J., Stenseth, N.C.S., 2004, Genetiske analyser påviser kompleks bestandsstruktur hos torsk i Skagerrak-Nordsjøområdet
Fisken og Havet, 2, 109-110