Ørjan Karlsen

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Telephone: 922 93 930

Curriculum Vitae

Stilling

Seniorforsker (forsker 1183)

Utdannelse

Cand. Scient. i 1992 ved  Institutt for Fiskeri- og Marinbiologi, Universitetet i Bergen, Norge. Tittel på oppgaven: Vekstmønster og ultralydestimert organutvikling gjennom første gytesesong hos torsk (Gadus morhua L.) i oppdrett. [Growth pattern and ultrasound based organ development through first spawning season in farmed cod (Gadus morhua L.)].

Dr. Scient. i 1999 ved Institutt for Fiskeri- og Marinbiologi, Universitetet i Bergen, Norge. Tittel på Dr. Scient. avhandlingen: Hatching and ontogeny in marine fishes: Consequences for variability and viability at first feeding

Publikasjoner (OrcID: orcid.org/0000-0001-5801-4200)

Halttunen, E., Gjelland, K.Ø., Hamel, S., Serra-Llinares, R.M., Nilsen, R., Arachevala-Lopez, P., Skardhamar, J., Johnsen, I.A., Asplin, L., Skaala, Ø., Karlsen, Ø., Bjørn, P.A. & Finstad, B. (2017). Sea trout adapt their migratory behaviour in response to high salmon lice concentrations. Journal of Fish Diseases

Vollset, K.W., Dohoo, I., Karlsen, Ø., Halttunen, E., Kvamme, B.O., Finstad, B., Wennevik, V., Diserud, O.H., Bateman, A., Friedland, K.D., Mahlum, S., Jørgensen, C., Qviller, L., Krkosek, M., Åtland, Å. & Barlaup, B.T. (2017). Food for thought: Disentangling the role of sea lice on the marine survival of Atlantic salmon. ICES Journal of Marine Science ( http://doi.org/10.1093/icesjms/fsx104 )

Sørensen, L., Sørhus, E., Nordtug, T., Incardona, J., Linbo, T., Giovanetti, L., Karlsen, Ø. & Meier, S. (2017). Oil droplet fouling and differential PAH toxicokinetics in embryos of Atlantic haddock and cod. PLOS ONE, e0180048 ( https://doi.org/10.1371/journal.pone.0180048 )

Vollset, K.W., Halttunen, E., Finstad, B., Bjørn, P.A. & Dohoo, I. (2017). Salmon lice infestations on sea trout predicts infestations on migrating salmon post-smolts. ICES Journal of Marine Science, ( http://doi.org/10.1093/icesjms/fsx090 )

Skjæraasen, J.E., Divine, J., Godiksen, J., Fonn, M., Otterå, H., Kjesbu, O.S., Norberg, B., Langangen, Ø. & Karlsen, Ø. (2017). Timecourse of oocyte development in saithe Pollachius virens. Journal of Fish Biology 90: 109-128. ( http://dx.doi.org/10.1111/jfb.13161 )

Sæle, Ø., Haugen, T., Karlsen, Ø., van der Meeren, T., Bæverfjord, G., Hamre, K., Rønnestad, I., Moren, M. & Lie, K.K. (2017). Ossification of Atlantic cod – developmental stages revisited. Aquaculture 468: 524-533. ( http://doi.dx.org/10.1016/j.aquaculture.2016.11.004 )

Karlsen, Ø., Amlund, H., Berg, A. & Olsen, R.E. (2017). The effect of dietary chitin on growth and nutrient digestibility in farmed Atlantic cod, Atlantic salmon and Atlantic halibut. Aquaculture Research 48: 123-133. ( http://dx.doi.org/10.1111/are.12867 )

Sørhus, E., Incardona, J.P., Karlsen, Ø., Linbo, T., Sørensen, L., Nordtug, T., van der Meeren, T., Thorsen, A., Thorbjørnsen, M., Jentoft, S., Edvardsen, R.B. & Meier, S. (2016). Crude oil exposures reveal roles for intracellular calcium cycling in haddock craniofacial and cardiac development. Scientific Reports 6: 31058. ( http://dx.doi.org/10.1038/srep31058 )

Valen, R., Eilertsen, M., Edvardsen, R.B., Furmanek, T., Rønnestad, I., van der Meeren, T., Karlsen, Ø., Nilsen, T. & Helvik, J.V. (2015). The two-step development of a duplex retina involves distinct events of cone and rod neurogenesis and differentiation. Developmental Biology 416: 389-406 ( http://dx.doi.org/10.1016/j.ydbio.2016.06.041 )

Otterå, H., Thorsen, A., Karlsen, Ø., Fjelldal, P.G., Morton, H.C. & Taranger, G.L.  (2016). Performance of triploid Atlantic cod (Gadus morhua L.) in commercial aquaculture. Aquaculture 464: 699-709. ( http://dx.doi.org/10.1016/j.aquaculture.2016.08.018 )

Bizuayehu, T.T., Furmanek, T., Karlsen, Ø., van der Meeren, T., Edvardsen, R.B., Rønnestad, I., Hamre, K., Johansen, S.D. & Babiak, I. (2016). First feed affects expression of microRNAs and their targets in Atlantic cod. British Journal of Nutrition, 115: 1145-1154. ( http://dx.doi.org/10.1017/S0007114516000155 )

Le-Hoang, T.M.D., Angotzi, R., Ebbesson, L., Karlsen, Ø. & Rønnestad, I. (2016). The ontogeny and brain distribution dynamics of the appetite regulators NPY, CART and pOX in larval Atlantic cod ( Gadus morhua L.). PLoS ONE 11(4): e015374 ( http://dx.doi.org/10.1371/journal.pone.0153743 )

Chauton, M.S., Galloway, T., Kjørsvik, E., Størseth, T.R., Puvanendran, V., van der Meeren, T., Karlsen, Ø., Rønnestad, I. & Hamre, K. (2015). 1H NMR metabolic profiling of cod (Gadus morhua) larvae: Potential effects of temperature and diet composition during early developmental stages. Biology Open 4: 1671-1678. ( http://dx.doi.org/10.1242/bio.014431 )

Penglase, S., Edvardsen, R.B., Furmanek, T., Rønnstad, I., Karlsen, Ø., van der Meeren, T. & Hamre, K. (2015). Diet affects the redox system in developing Atlantic cod (Gadus morhua) larvae. RedOx Biology 5: 308-318. ( http://dx.doi.org/10.1016/j.redox.2015.06.003 )

Duan, Y., Stien, L.H., Thorsen, A., Karlsen, Ø., Sandlund, N., Li, D., Fu, Z. & Meier, S. (2015). An automatic counting system for transparent pelagic fish eggs based on computer vision. Aquaculture Engineering 67: 8-13. ( http://dx.doi.org/10.1016/j.aquaeng.2015.05.001 )

Karlsen, Ø., van der Meeren, T., Rønnestad, I., Mangor-Jensen, A., Galloway, T.F., Kjørsvik, E. & Hamre, K. (2015). Copepods enhance nutritional status, growth, and development in Atlantic cod (Gadus morhua L.) larvae – can we identify the underlying factors? PeerJ, 3: e902. ( http://dx.doi.org/10.7717/peerj.902 )

Sørhus, E., Edvardsen, R.B., Karlsen, Ø., Nordtug, T., van der Meeren, T., Thorsen, A., Harman, C., Jentoft, S., Meier, S. (2015). Unexpected interaction with dispersed crude oil droplets drives severe toxicity in Atlantic haddock embryos. PLOS ONE 10(4): e0124376. ( http://dx.doi.org/10.1371/journal.pone.0124376 )

Taranger, G.L., Karlsen, Ø., Bannister, R.J., Glover, K.A., Husa, V., Karlsbakk, E., Kvamme, B.O., Boxaspen, K.K., Bjørn, P.A., Finstad, B., Madhun, A.S., Morton, H.C. & Svåsand, T. (2015). Risk assessment of the environmental impact of Norwegian Atlantic salmon farming. ICES Journal of Marine Sciences 72: 997-1021. ( http://dx.doi.org/10.1093/icesjms/fsu132 )

van der Meeren, T., Karlsen, Ø., Liebig, P.L. & Mangor-Jensen, A. (2014). Copepod production in a saltwater pond system: a reliable method for achievement of natural prey in start-feeding of marine fish larvae. Aquacultural Engineering 62: 17-27. ( http://dx.doi.org/10.1016/j.aquaeng.2014.07.003 )

Uglem, I., Karlsen, Ø., Sanchez-Jerez, P. & Sæther, B.-S. (2014). Impacts of attraction of wild marine fish to open cage salmonid farms in Norway. Aquaculture Environment Interactions 6: 91-103. ( http://dx.doi.org/10.3354/aei00112 )

Karlsen Ø., Mittelholzer, C., Andersson, E., Norberg, B. and Taranger G.L. (2014). Continuous light affects onset of puberty and associated changes in pituitary gonadotropin subunit transcript levels, and plasma estradiol-17β and testosterone levels in Atlantic cod (Gadus morhua L.) females. Aquaculture 425-425: 95-103. ( http://dx.doi.org/10.1016/j.aquaculture.2013.12.029 )

Zhou, Z., Karlsen, Ø., He, S., Olsen, R.E., Yao, B. & Ringø, E. (2013). The effect of dietary chitin on the autochthonous gut bacteria of Atlantic cod (Gadus morhua L.). Aquaculture Research 44: 1889-1900. ( http://dx.doi.org/10.1111/j.1365-2109.2012.03194.x )

Karlsen, Ø., Otterå. H., Thorsen, A. (2013). Fertility of gynogenetic Atlantic cod (Gadus morhua L.). Journal of Applied Ichthyology 29: 1292-1296. ( http://dx.doi.org/10.1111/jai.12272 )

Skulstad, O.F., Taylor, J., Davie, A., Kristiansen, T., Mayer, I., Migaud, H., Taranger, G.L., Olsen, R.E. & Karlsen, Ø. (2013). Effects of light regime on diurnal plasma melatonin levels and vertical distribution in farmed Atlantic cod (Gadus morhua L.). Aquaculture 414-415: 280-287. ( http://dx.doi.org/10.1016/j.aquaculture.2013.08.007 )

Kleppe, L., Karlsen, Ø., Edvardsen, R.B., Norberg, B., Andersson, E., Furmanek, T., Taranger, G. L. & Wargelius, A. (2013). Cortisol treatment of prespawning female cod affects cytogenesis related factors in eggs and embryos. General and Comparative Endocrinology 189: 84-95. ( http://dx.doi.org/10.1016/j.ygcen.2013.04.028 )

Korsøen, Ø.J., Dempster, T., Fosseidengen, J.E., Karlsen, Ø., Oppedal, F., Stien, L.H. & Kristiansen, T. (2013). Towards cod without spawning: artificial continuous light in submerged sea-cages maintains growth and delays sexual maturation for farmed Atlantic cod (Gadus morhua). Aquaculture Environment Interactions 3: 245-255. ( http://dx.doi.org/10.3354/aei00067 )

Hansen, A-C., Rosenlund, G., Karlsen, Ø., Olsen, R.E. & Hemre, G-I. (2013). Marine ash-products influence growth, feed utilization and liver size when Atlantic cod Gadus morhua L. are fed plant based diets. Journal of Applied Ichthyology 29: 532-540. ( http://dx.doi.org/10.1111/jai.12087 )

Skulstad, O.F., Karlsen, Ø., Fosseidengen, J.E., Taranger, G.L. & Oppedal, F. (2013). Vertical distribution and sexual maturation in cage-farming of Atlantic cod (Gadus morhua L.) exposed to natural or continuous light. Aquaculture Research 44: 903-917. ( http://dx.doi.org/10.1111/j.1365-2109.2012.03095.x )

Opstad, I., Fjelldal, P.G., Karlsen, Ø., Thorsen, A., Hansen, T.J. & Taranger, G.L. (2013). The effect of triploidization of Atlantic cod (Gadus morhua L.); survival, growth and deformities during early life stages. Aquaculture 388: 54-59. ( http://dx.doi.org/10.1016/j.aquaculture.2013.01.015 )

Meager, J.J., Skjæraasen, J.E., Karlsen, Ø., Løkkeborg, S., Mayer, I., Michalsen, K., Nilsen, T. & Fernö, A. (2012). Environmental regulation of individual depth on a cod spawning ground. Aquatic Biology 17: 211-221. ( http://dx.doi.org/10.3354/ab00469 )

Bogevik, A.S., Natário, S., Karlsen, Ø., Thorsen, A., Hamre, K., Rosenlund, G. and Norberg, B. (2012). The effect of dietary lipid content and stress on egg quality in farmed Atlantic cod (Gadus morhua L.). Journal of Fish Biology 81: 1391-1405. ( http://dx.doi.org/10.1111/j.1095-8649.2012.03425.x )

Otterå, H., Thorsen, A., Peruzzi, S., Dahle, G., Hansen, T & Karlsen, Ø. (2011). Induction of meiotic gynogenesis in Atlantic cod, Gadus morhua (L.). Journal of Applied Ichthyology 27: 1298-1302. ( http://dx.doi.org/10.1111/j.1439-0426.2011.01863.x )

Almeida, F.F.L., Andersson, E., Mittelholzer, C., Karlsen, Ø., Taranger, G.L. & Schulz, R.W. (2011). Pituitary gonadotropin and testicular gonadotropin receptor expression in Atlantic cod (Gadus morhua, L.) during the first reproductive season: Effects of photoperiod modulation. General and Comparative Endocrinology 173: 111-11 ( http://dx.doi.org/10.1016/j.ygcen.2011.05.002 )

Skjæraasen, J.E., Meager, J.J., Karlsen, Ø., Hutchings, J.A., Fernö, A. (2011). Extreme spawning site fidelity in Atlantic cod. ICES Journal of Marine Science 68: 1472-1477. ( http://dx.doi.org/10.1093/icesjms/fsr055 )

Hansen, A.-C., Hemre, G.-I., Karlsen, Ø., Koppe, W., Rosenlund, G. (2011). Do plant based diets for Atlantic cod (Gadus morhua L.) need additions of crystalline lysine or methionine? Aquaculture Nutrition 17: 362-371. ( http://dx.doi.org/10.1111/j.1365-2095.2010.00770.x )

Skjæraasen, J.E., Meager, J.J., Karlsen, Ø., Mayer, I., Dahle, G, Rudolfsen, G. & Fernö, A. (2010). Mating competition between farmed and wild cod, Gadus morhua. Marine Ecology Progress Series 412: 247-258. ( http://dx.doi.org/10.3354/meps08670 )

Lekva, A., Hansen, A.-C., Rosenlund, G., Karlsen, Ø. Hemre, G.-I. (2010). Energy dilution with α-cellulose in diets for Atlantic cod (Gadus morhua L.) juveniles - Effects on growth, feed intake, liver size and digestibility of nutrients. Aquaculture 300: 169-175. ( http://dx.doi.org/10.1016/j.aquaculture.2010.01.001 )

Taranger, G.L., Carrillo, M., Schulz, R.W., Fontaine, P., Zanuy, S., Felip, A., Weltzien, F.-A., Dufour, S., Karlsen, Ø., Norberg, B., Andersson, E. & Hansen, T. (2010). Control of puberty in farmed fish. General and Comparative Endocrinology 165: 483-515. ( http://dx.doi.org/10.1016/j.ygcen.2009.05.004 )

Meager, J.J., Skjæraasen, J.E., Fernö, A. & Karlsen, Ø., Løkkeborg, S., Michalsen, K. & Utskot, S.O. (2009). Vertical dynamics and reproductive behaviour of farmed and wild Atlantic cod, Gadus morhua. Marine Ecology Progress Series 389: 233-243. ( http://dx.doi.org/10.3354/meps08156 )

Otterå, H.O., Slinde, E., Olsen, R.E. & Karlsen, Ø. (2009). Quality of wild captured saithe (Pollachius virens L.) fed formulated diets for eight months. Aquaculture Research 40: 1310-1319. ( http://dx.doi.org/10.1111/j.1365-2109.2009.02230.x )

Mittelholzer, C., Andersson, E., Taranger, G.L., Karlsen, Ø., Norberg, B. (2009). Quantification of gonadotropin subunits GPα, FSHβ, and LHβ mRNA expression from Atlantic cod (Gadus morhua) throughout a reproductive cycle. Comparative Biochemistry and Physiology. Part B 153: 288-295. (http://dx.doi.org/10.1016/j.cbpb.2009.03.011) http://dx.doi.org/10.1016/j.cbpb.2009.03.011

Almeida, F.F.L., Taranger, G.L., Norberg, B., Karlsen, Ø., Bogerd, J. & Schulz, R.W. (2009). Photoperiod-modulated testis maturation in Atlantic cod (Gadus morhua, L.). Biology of Reproduction 80: 631-640. ( http://dx.doi.org/10.1095/biolreprod.108.071340 )

Skjæraasen, J.E., Mayer, I., Meager, J.J., Rudolfsen, G., Karlsen, Ø., Haugland, T. & Kleven, O. (2009). Sperm characteristics and competitive ability in farmed and wild cod. Marine Ecology Progress Series 375: 219-228. ( http://dx.doi.org/10.3354/meps07774 )

Skjæraasen, J.E., Meager, J.J. & Karlsen, Ø. (2008). The expression of secondary sexual characteristics in recruit- and repeat-spawning farmed and wild Atlantic cod (Gadus morhua). ICES Journal of Marine Sciences 65: 1710-1716. ( http://dx.doi.org/10.1093/icesjms/fsn147 )

Olsen, R.E., Sundell, K., Ringø, E., Myklebust, R., Hemre, G.-I., Hansen, T. & Karlsen, Ø. (2008). The acute stress response in fed and food deprived Atlantic cod, Gadus morhua L. Aquaculture 280: 232-241. ( http://dx.doi.org/10.1016/j.aquaculture.2008.05.006 )

Olsen, R.E., Hansen A.-C., Rosenlund, G., Hemre, G.-I., Mayhew, T.M., Knudsen, D.L., Eroldoðan, O.T., Myklebust, R. & Karlsen, Ø. (2007). Total replacement of fish meal with plant proteins in diets for Atlantic cod (Gadus morhua L.) – II Health aspects. Aquaculture 272: 612-624. ( http://dx.doi.org/10.1016/j.aquaculture.2007.05.010 )

Hansen, A.-C., Rosenlund, G., Karlsen, Ø., Koppe, W. & Hemre, G.-I. (2007). Total replacement of fish meal with plant proteins in diets for Atlantic cod (Gadus morhua L.) I - Effects on growth and protein retention. Aquaculture 272: 599-611. ( http://dx.doi.org/10.1016/j.aquaculture.2007.08.034 )

Moren, M., Malde, M.K., Olsen, R.E., Hemre, G.-I., Dahl, L., Karlsen, Ø. & Julshamn, K. (2007). Fluorine accumulation in Atlantic salmon (Salmo salar), Atlantic cod (Gadus morhua), rainbow trout (Onchorhyncus mykiss) and Atlantic Halibut (Hippoglossus hippoglossus)  fed diets with krill or amphipod meals and fish meal based diets diets with sodium fluoride (NaF) inclusion. Aquaculture 269: 525-531. ( http://dx.doi.org/10.1016/j.aquaculture.2007.04.059 )

Suontama, J., Karlsen, Ø., Moren, M., Hemre, G.-I., Melle, W., Langmyhr, E., Mundheim, H., Ringø, E. & Olsen. R.E. (2007). Growth, feed conversion and chemical composition of Atlantic salmon (Salmo salar L.) and Atlantic halibut (Hippoglossus hippoglossus L.) fed diets supplemented with krill or amphipods. Aquaculture Nutrition 13: 241-255. ( http://dx.doi.org/10.1111/j.1365-2095.2007.00466.x )

Otterå, H., Carlehög, M., Karlsen, Ø., Akse, L., Borthen, J. & Eilertsen, G. (2007). Effect of diet and season on quality of farmed Atlantic cod (Gadus morhua L). LWT – Food Science and Technology 40: 1623-1629. ( http://dx.doi.org/10.1016/j.lwt.2006.09.003 )

Hansen, A.-C., Karlsen, Ø., Rosenlund, G., Rimbach, M. & Hemre, G.-I. (2007). Dietary plant-protein utilisation in Atlantic cod, Gadus morhua. Aquaculture Nutrition 13: 200-215. ( http://dx.doi.org/10.1111/j.1365-2095.2007.00486.x )

Karlsen, Ø., Sountama, J. & Olsen. R.E. (2006). Effect of Antarctic krillmeal on quality of farmed Atlantic cod (Gadus morhua L.). Aquaculture Research 37: 1676-1684. ( http://dx.doi.org/10.1111/j.1365-2109.2006.01615.x )

Moren, M., Suontama, J., Hemre, G.-I., Karlsen, Ø., Olsen, R.E., Mundheim, H. & Julshamn, K. (2006). Element concentrations in meals from krill and amphipods, - possible alternative protein sources in complete diets for farmed fish. Aquaculture 261: 174-181. ( http://dx.doi.org/10.1016/j.aquaculture.2006.06.022 )

Hansen, A.-C., Rosenlund, G., Karlsen, Ø., Olsvik, P. & Hemre, G.-I. (2006). The inclusion of plant protein in cod diets, its effects on macronutrient digestibility, gut and liver histology and heat shock protein transcription. Aquaculture Research 37: 773-784. ( http://dx.doi.org/10.1111/j.1365-2109.2006.01490.x )

Karlsen, Ø., Hemre, G.-I., Tveit, K. & Rosenlund, G. (2006). Effect of varying levels of macro nutrients and continuous light on growth, energy deposits and maturation in farmed Atlantic cod (Gadus morhua L.). Aquaculture 255: 242-254. ( http://dx.doi.org/10.1016/j.aquaculture.2005.12.029 )

Jørstad, K.E., Karlsen, Ø., Svåsand, T. and Otterå, H. (2006). Comparison of growth rate among different protein genotypes in Atlantic cod, Gadus morhua, under farmed conditions. ICES Journal of Marine Sciences 63: 235-245. ( http://dx.doi.org/10.1016/j.icesjms.2005.11.005 )

Karlsen, Ø., Norberg, B., Kjesbu, O.S. and Taranger, G.L. (2006). Effects of photoperiod and exercise on growth, liver size, and age at puberty in farmed Atlantic cod (Gadus morhua L.). ICES Journal of Marine Sciences 63: 355-364. ( http://dx.doi.org/10.1016/j.icesjms.2005.10.013 )

Stien, L.H., Hirmas, E., Bjørnevik, M., Karlsen, Ø., Nortvedt, R., Rørå, A.M.B., Sunde, J. & Kiessling, A. (2005). The effects of stress and storage temperature on the colour and texture of pre-rigor filleted farmed cod (Gadus morhua L.). Aquaculture Research 36: 1197-1206. ( http://dx.doi.org/10.1111/j.1365-2109.2005.01339.x )

Skjæraasen, J.E., Salvanes, A.G., Karlsen, Ø., Dahle, R., Nilsen, T. & Norberg, B. (2004). The effect of photoperiod on sexual maturation, appetite and growth in wild Atlantic cod (Gadus morhua). Fish Physiology and Biochemistry 30: 163-174. ( http://dx.doi.org/10.1007/s10695-005-4319-6 )

Rosenlund, G., Karlsen, Ø., Tveit, K., Mangor-Jensen, A & Hemre, G.-I. (2004). Effect of feed composition and feeding frequency on growth, feed utilisation and nutrient retention in juvenile cod, Gadus morhua. Aquaculture Nutrition 10: 371-378. ( http://dx.doi.org/10.1111/j.1365-2095.2004.00312.x )

Knudsen, F.R., Fosseidengen, J.E., Oppedal, F., Karlsen, Ø. & Ona, E. (2004). Hydroacoustic monitoring of fish in sea cages: target strength (TS) measurements on Atlantic salmon (Salmo salar). Fisheries Research 69: 205-209. ( http://dx.doi.org/10.1016/j.fishres.2004.05.008 )

Hemre, G.-I., Karlsen, Ø., Eckhoff, K., Tveit, K., Mangor-Jensen, A. & Rosenlund G. (2004). Effect of season, light regime and diet on muscle composition and selected quality parameters in farmed Atlantic cod, Gadus morhua L. Aquaculture Research 35: 683-697. ( http://dx.doi.org/10.1111/j.1365-2109.2004.01069.x )

Høie, H., Andersson, C.A., Folkvord, A. & Karlsen, Ø. (2004). Precision and accuracy of stable isotope signals in otoliths of pen-reared cod (Gadus morhua L.) when sampled with high-resolution micromill. Marine Biology 144: 1039-1049. ( http://dx.doi.org/10.1007/s00227-003-1275-5 )

Weltzien, F.-A., Karlsen, Ø., Norberg, B., 2003. Growth patterns and plasma levels of testoterone, 11-ketotestosterone, and IGF-1 in male Atlantic halibut (Hippoglossus hippoglossus) from juvenile stages throughout sexual development. Fish Physiology and Biochemistry 28: 227-228. ( http://dx.doi.org/10.1023/B:FISH.0000030539.22672.fc )

Dahle, R., Taranger, G.L., Karlsen, Ø., Kjesbu, O.S. & Norberg, B. (2003). Gonadal development and associated changes in relative liver size and sexual steroids during the reproductive cycle of captive male and female Atlantic cod (Gadus morhua L.). Comparative Biochemistry and Physiology part A 136: 641-653. ( http://dx.doi.org/10.1016/S1095-6433(03)00215-0 )

Bjørnevik, M., Karlsen, Ø., Johnston, I. & Kiessling, A. (2003). Effect of sustained exercise on white muscle structure and flesh quality in farmed cod (Gadus morhua L.). Aquaculture Research 34: 55-64. ( http://dx.doi.org/10.1046/j.1365-2109.2003.00794.x )

Hemre, G.-I., Karlsen, Ø., Mangor-Jensen, A. & Rosenlund, G. (2003). Digestibility of dry matter, protein and lipid by cod, Gadus morhua, comparison of sampling methods. Aquaculture 225: 225-232. ( http://dx.doi.org/10.1016/S0044-8486(03)00291-6 )

Weltzien, F.-A., Taranger, G.L., Karlsen, Ø. & Norberg, B. (2002). Spermatogenesis and related plasma androgen levels in Atlantic halibut (Hippoglossus hippoglossus L.). Comparative Biochemistry and Physiology part A 132: 567-575. ( http://dx.doi.org/10.1016/S1095-6433(02)00092-2 )

Hansen, T., Karlsen, Ø., Taranger, G.L., Hemre, G.-I., Holm, J.C. & Kjesbu, O.S. (2001). Growth, gonadal development and spawning time of Atlantic cod (Gadus morhua) reared under different photoperiods. Aquaculture 203: 51-67 ( http://dx.doi.org/10.1016/S0044-8486(01)00610-X )

Karlsen, Ø. & Mangor-Jensen, A. (2001). A correlation between phototactic response and first-feeding of (Hippoglossus hippoglossus L.) larvae. Aquaculture Research 32: 907-912. ( http://dx.doi.org/10.1046/j.1365-2109.2001.00628.x )

Norberg, B., Weltzien, F.-A., Karlsen, Ø. & Holm, J.C. (2001). Effects of photoperiod on sexual maturation and somatic growth in male Atlantic halibut (Hippoglossus hippoglossus L.). Comparative Biochemistry and Physiology part B 129: 357-365. ( http://dx.doi.org/10.1016/s1096-4959(01)00320-7 )

Porter, M.J.R., Stefansson, S.O., Nyhammer, G., Karlsen, Ø., Norberg, B. & Bromage, N.R. (2000). Environmental influences on melatonin secretion in Atlantic cod (Gadus morhua L.) and their relevance to commercial culture. Fish Physiology and Biochemistry 23: 191-200. ( http://dx.doi.org/10.1023/a:1007816426509 )

Karlsen, Ø. Taranger, G.L., Dahle, R. & Norberg, B. (1999). Effects of exercise and continuous light on early sexual maturation in farmed Atlantic cod (Gadus morhua L.). Proceedings from the 6th International Symposium on Reproductive Physiology of Fish, Bergen, Norway, July 4-9, 1999. Pp. 328-330.

Karlsen, Ø., Skiftesvik, A.B. & Helvik, J.V. (1998). The effect of light on activity and growth of Atlantic halibut, Hippoglossus hippoglossus L., yolk-sac larvae. Aquaculture Research 29: 899-912. ( http://dx.doi.org/10.1046/j.1365-2109.1998.29120899.x )

Helvik, J.V. & Karlsen, Ø. (1996). The effect of light- and dark-rearing on the development of the eyes of Atlantic halibut (Hippoglossus hippoglossus) yolk sac larvae. Marine and Freshwater Behaviour and Physiology 28: 107-121. ( http://dx.doi.org/10.1080/10236249609378982 )

Karlsen, Ø., Holm, J.C. & Kjesbu, O.S. (1995). Effects of periodic starvation on reproductive investment in first-time spawning Atlantic cod (Gadus morhua L.). Aquaculture 133: 159-170. ( http://dx.doi.org/10.1016/0044-8486(94)00399-9 )

Karlsen, Ø. & Holm, J.C. (1994). Ultrasonography, a non-invasive method for sex determination in cod (Gadus morhua L.). Journal of Fish Biology 44: 965-971. ( http://dx.doi.org/10.1006/jfbi.1994.1089 )

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