Knut Yngve Børsheim

Telephone: 97781312
Department: Oseanografi og klima

Arbeider med

Mikrobielle prosesser i marin karbonsyklus og  biogeokjemiske påvirkninger av utvekslingen av karbondioksid mellom hav og atmosfære. Sentralt står bakterienes aktivitet og omdanning av løst organisk materiale samt global karbonsyklus og forsuring av havet.

Curriculum Vitae


Institute of Marine Research
Nordnesgaten 33
NO-5017 Bergen, Norway
Phone: +47 55238491
Age and nationality
Date of birth: Feb. 24 1953
Nationality: Norwegian
Home address: Haugeveien 3, NO-5005 Bergen

Academic degrees
Doctor philosophiae (University of Bergen, Norway, 1993).
Title of the thesis for the dr. degree: “Growth rate and mortality of bacteria in aquatic environments”. Opponents: Professor Morten Søndergaard, University of Copenhagen, Denmark, and Professor Åke Hagström, University of Umeå, Sweden.
Cand. real.(University of Bergen, Norway 1979). Title of the thesis: “Carbon and Sulfur Cycles in Lake Sælenvatnet” (In Norwegian).

Present affiliations
Institute of Marine Research, Climate and Oceanography Group, Bergen, Norway
Bjerknes Centre for Climate Research, Bergen, Norway

Professional history
Scientific Assistant 1978-1979 at: Department of Microbiology and Plant Physiology, The University of Bergen, Norway.
Scientific Assistant 1980-1982 at: Department of Microbiology, the Norwegian Agricultural University, Ås, Norway.
Research scientist at 1983-1989: Department of Microbiology and Plant Physiology, The University of Bergen, Norway.
Senior Research Scientist 1990-2004: (20-30% of the time also Associate Professor) at the Department of Biotechnology, Norwegian University of Science and Technology, Trondheim, Norway.
Senior Research Scientist 2005-2008 (20%) at Institute of Biology, University of Bergen).
Senior Research Scientist 2005 –current: Institute of Marine Research, Bergen, Norway

Publications in peer reviewed scientific periodicals

Børsheim, K.Y., Milutinivic, S., Drinkwater, K.F. 2014. TOC and satellite-sensed chlorophyll and primary production at the Arctic Front in the Nordic Seas. Journal of Marine Systems, 139:373-382.
Skogen, M.D., Olsen, A., Børsheim, K.Y., Sandø, A.B., Skjelvan, I. 2014. Modelling ocean acidification in the Nordic and Barents Seas in present and future climate. Journal of Marine Systems, 131: 10-20.
Børsheim, K.Y., Drinkwater, K.F. 2014 Different temperature adaptation in Arctic and Atlantic heterotrophic bacteria in the Barents Sea Polar Front region. Journal of Marine Systems, 130: 160-166.
Harrison, W.G. Børsheim, K.Y., Li, W.K.W., Maillet, G.L., Pepin, P., Sakshaug, E., Skogen, M.D., Yeats, P.A.  2013. Phytoplankton production and growth regulation in the Subarctic North Atlantic: A comparative study of the Labrador Sea-Labrador/Newfoundland shelves and Barents/Norwegian/Greenland seas and shelves. Progress in Oceanography, 114: 26-45.
Thingstad T.F., R.G.J. Bellerby, G. Bratbak, K.Y. Børsheim, J.K. Egge, M. Heldal, A. Larsen, C. Neill, J. Nejstgaard, S. Norland, R.‐A. Sandaa, E.F. Skjoldal, T. Tanaka, R. Thyrhaug, B. Töpper (2008) Counterintuitive carbon‐to‐nutrient coupling in an Arctic pelagic ecosystem Nature 455, 387 – 390
Myklestad, S.M., Børsheim, K.Y. (2007) Dynamics of carbohydrates in the Norwegian Sea
inferred from monthly profiles collected during 3 years at 66⁰N2⁰E. Marine Chemistry 107:475‐485
Ballance., S., Børsheim, K.Y., Inngjerdingen, K., Smestad Paulsen, B., Christensen, B.E. (2007) A re-examination and partial characterisation of polysaccharides released by mild acid hydrolysis from the chlorite-treated leaves of Sphagnum papillosum. Carbohydrate Polymers 67:104-115
Børsheim, K.Y., Vadstein, O., Myklestad, S.M., Reinertsen, H., Kirkvold, S., Olsen, Y. (2005) Photosynthetic algal production, accumulation and release of phytoplankton storage carbohydrates, and bacterial production in a gradient in daily nutrient supply. Journal of Plankton Research 47:743-755
Ballance, S., Børsheim, Y., Christensen, B.E. (2003) A unique 5-keto sugar in the leaves of Sphagnum moss: A review of the evidence. In: J. Päivänen (Ed.).12th International Peat Congress: 293-299. Finnish Peatland Society, Tampere, Finland.
Legrand, C., Johansson, N., Johnsen, G., Børsheim, K.Y., Granéli, E. (2001) Phagotrophy and toxicity variation in the mixotrophic Prymnesium patelliferum (Haptophyceae). Limnology and Oceanography 46:1208-1214
Heggestad, T., Børsheim, K.Y. (2002) The accessibility of and distribution of the Norwegian National Air Emergency  Service. Air Medical Journal 21:39-45
Børsheim, K.Y., Christensen, B.E., Painter, T.J. (2001) Preservation of fish by embedment in Sphagnum moss, peat, or holocellulose: experimental proof of the oxopolysaccharidic nature of the preservative substance and its antimicrobial and tanning action. Innovative Food Science and Emerging Technologies  2:63-74
Børsheim, K.Y (2000) Bacterial production rates and concentrations of organic carbon at the end of the growing season in the Greenland Sea. Aquatic Microbial Ecology 21:115-123
Noji, T.M., Børsheim, K.Y., Rey, F., Nortvedt, R (1999) Dissolved organic carbon associated with sinking particles can be crucial for estimates of vertical carbon flux. Sarcia 84:129-135
Noji, T., Miller, L., Skjelvan, I., Falck, E., Børsheim, K.Y., Rey, F., Urban-Rich, J., Johannessen, T (1999) Constraints on carbon drawdown in the Greenland Sea. In: Schäfer, P., Ritzrau, W, Schlüter, M., Thiede, J. (eds). The Northern North Atlantic. A changing environment. Springer, Berlin.
Børsheim, K.Y., Myklestad, S.M., Sneli, J-.A. (1999). Monthly profiles of DOC, mono- and polysaccharides in the Trondheimsfjord (Norway) during two years. Marine  Chemistry 63:255-272
Børsheim, K.Y., Myklestad, S.M. (1997) Dynamics of DOC in the Norwegian Sea inferred from monthly profiles collected during three years at 66oN2oE. Deep-Sea Research 44:593-601
Noji, T.T., Rey, F., Miller, L., Børsheim, K.Y., Hirche, H.-J., Urban-Rich, J. (1996). Plankton dynamics and sedimentation. In: Wadams, P., Wilkinson, J.P. and Wells, S.C.S (eds.) European Subpolar Ocean Programme, Sea Ice - Ocean Interactions. University of Cambridge, U.K.
Turley, C.M., Børsheim, K., Iriberri, J., Prosser, J (1996) The estimation of bacterial biomass in Mediterranean seawater. Handbook of Method Protocols. Quality Assurance Pilot Study for the Mediterranean Targeted project. The European Commssion, Brussels.
Børsheim, K.Y. (1996). The seasonal and spatial variation of dissolved organic carbon in sea water. In: Wadams, P., Wilkinson, J.P. and Wells, S.C.S (eds.) European Subpolar Ocean Programme, Sea Ice - Ocean Interactions. University of Cambridge, U.K.
Hitchcock, G. L., Rossby., T., Lillibridge III, J. L., Lessard, E. J., Levine, E. R., Connors, D. N., Børsheim, K. Y., Mork, M. (1994). Signatures of stirring and mixing near the Gulf Stream front. J. Mar. Res. 52:797-836
Børsheim, K.Y. (1993). Native marine bacteriophages. FEMS Microbiology Ecology 102:141-159
Børsheim, K.Y. (1992). Growth rate and mortality of bacteria in aquatic environments. Thesis for the degree Dr. philos. at the University of Bergen.
Putt, M., Børsheim, K. Y., Miceli, G., Stoecker, D. K. (1991). Seasonal changes in cell size and abundance of bacterioplankton during the Phaeocystis sp. bloom in McMurdo Sound. Antarctic J. of the United States 1991:199-20
Børsheim, K.Y. (1990). Bacterial biomass and production rate in the Gulf Stream front regions. Deep-Sea Research 37:1297-1309
Børsheim, K.Y., Bratbak., G., Heldal, M. (1990). Enumeration and biomass estimation of planktonic bacteria and viruses by transmission electron microscopy. Appl. Environ. Microbiol. 56:352-356
Bergh, Ø., Børsheim, K.Y., Bratbak., G., Heldal, M. (1989). High abundance of viruses found in aquatic environments. Nature 340:467-468
Børsheim, K.Y., Harboe, T., Johnsen, T., Norland, S., Nygaard K. (1989) Flow cytometric characterization and enumeration of Chrysochromulina polylepis (Manton & Parke) during a bloom along the Norwegian coast. Mar. Ecol. Prog. Ser. 54:307-309
Andersen, S., Johnsen, G., Børsheim, K.Y. (1989). Changes in seasonal successions in Lake Kvernavatnet, compared to the PEG model. Arch. Hydrobiol. 115:401-415
Børsheim, K.Y., Norland, S. (1989). A counting device for oyster larvae. Aquaculture. 78:379-382
Børsheim, K.Y., Andersen, S., Johnsen, G., Kambestad, E.Ø., Norland, S. (1988). Primary and bacterial production compared to growth and food requirements of Daphnia longispina in Lake Kvernavatnet, West Norway. J. Plankton Res. 10:921-939
Mortensen, S.H., Børsheim, K.Y., Rainuzzo, J.R., Knutsen, G. (1988). Fatty acid and elemental composition of the marine diatom Chaetoceros gracilis Schütt. Effects of silicate deprivation, temperature and light intensity. J. Exp. Mar. Biol. Ecol. 122:173-185
Johnsen, G.H., Børsheim, K.Y. (1988). Functional response and food selection of the water flea, Bosmina longispina. J. Plankton Res. 10:319-325
Nygaard, K., Børsheim, K.Y., Thingstad, T.F. (1988). Grazing rates on bacteria by marine heterotrophic microflagellates compared to uptake rates of bacterial-sized, monodisperse fluorescent latex beads. Mar. Ecol. Prog. Ser. 44:154-165
Børsheim, K.Y., Bratbak, G. (1987). Cell volume to cell carbon conversion factors for a bacterivorous Monas sp. enriched from seawater. Mar. Ecol. Prog. Ser. 36:171-175
Børsheim, K.Y., Andersen, S. (1987). Grazing and food size selection by crustacean zooplankton compared to production of bacteria and phytoplankton in a shallow Norwegian mountain lake. J. Plankton Res. 9:367-369
Olsen, Y., Jensen, A., Reinertsen, H., Børsheim, K.Y., Heldal, M., Langeland, A. (1986). Dependence of the rate of release of phosphorus by zooplankton on the P:C ratio in the food supply, as calculated by a recycling model. Limnol. Oceanogr. 31:34-44
Børsheim, K.Y., Kuenen, J.G., Gottschal, J., Dundas, I.E.D. (1985). Microbial activities and chemical gradients in relation to the precicion of the sampling procedure. FEMS Microbiol. Ecol. 31:337-350
Larsson, P., Andersen, S., Børsheim, K.Y., Jakobsen, O., Johnsen, G. (1985). Individual growth of Daphnia longispina in the summer decline of the population. Arch. Hydrobio. Beih. Ergebn. Limnol. 21:341-350
Børsheim, K.Y., Olsen, Y. (1984). Grazing activities by Daphnia pulex on natural populations of bacteria and algae. Verh. Internat. Ver. Limnol. 22:644-648
Børsheim, K.Y. (1984). Clearance rate of bacteria-sized particles by freshwater ciliates, measured with monodisperse, fluorescent latex beads. Oecologia 63:286-288

Other publications

Børsheim, K.Y. (2008). Viruses. In Liam, F., Brewer, P. (eds.) Changes in surface CO2 and ocean pH in ICES shelf sea ecosystems. ICES Cooperative Research Report No. 280.
Børsheim, K.Y., Golmen, L (2010) Forsuring av havet. Kunnskapsstatus for norske farvann. SFT rapport TA nr 2575/2010
Y. Børsheim, K. Sokolov, O. Tito: Bacteria and viruses. In: Stiansen, J.E., Korneev, O., Titov, O., Arneberg, P. (Eds.), Filin, A., Hansen, J.R., Høines, Å., Marasaev, S. (Co-eds.) 2009. Joint Norwegian-Russian environmental status 2008. Report on the Barents Sea Ecosystem. Part II – Complete report. IMR/PINRO Joint Report Series, 2009(3), 375 pp. ISSN 1502-8828. (pp 30-33)
Børsheim Knut Yngve, 2012. Ocean acidification - A Norwegian perspective. IMBER newsletter, Issue n°21 - September 2012, Chapter 4.4


The John Martin Award from American Society of Limnology and Oceanography 2011 for the paper Bergh et al. (1989). The John Martin Award, established in 2005, recognizes a paper in aquatic sciences that is judged to have had a high impact on subsequent research in the field. The Martin Award is for papers at least 10 years old.


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